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Multiple realities
(covers information from several alternate timelines)
For the general concept of a department, please see Department.

Aboard Federation starships, ship duties among the Starfleet crew were delegated to individual departments, each headed by a department head. (TOS: "Where No Man Has Gone Before") These included science departments. (TOS: "Operation -- Annihilate!"; TNG: "The Game") Each department contained members of respective Starfleet divisions.

After Charles Tucker became chief engineer of Columbia, Captain Erika Hernandez received two transfer requests from his former department aboard Enterprise. (ENT: "Affliction")

In the alternate reality James T. Kirk, after becoming acting captain, ordered all departments at battle stations and ready in ten minutes. (Star Trek)

In 2266, Captain James T. Kirk of the USS Enterprise requested of Leonard McCoy that he find someone in his department with psychiatric and penology experience, if possible. (TOS: "Dagger of the Mind")

He might have meant the Life sciences or Medical department, as the former seemed headed by the Chief Medical Officer and the latter contained medical personnel.

After the Enterprise encountered a black star in 2267, Captain Kirk requested that all departments tie in with the record computer. (TOS: "Tomorrow is Yesterday")

In 2267, faced with anomalous aging at the Gamma Hydra IV, Kirk became concerned about Romulan involvement and in a staff meeting asked everyone to check out everything in their own departments, no matter how remote or far-fetched. (TOS: "The Deadly Years")

Aboard the USS Enterprise-D, Captain Picard, or William Riker often requested all departments to prepare reports, or to ready themselves for an impending crisis. In 2364, Picard expressed his wish to have all departments prepared for a warp 6 trip into the Neutral Zone. (TNG: "Angel One") In 2370, after a drill, Riker requested all departments to submit drill evaluation reports. (TNG: "Lower Decks") The same year, Picard requested a battle readiness report from all departments in response to what he believed was a potentially dangerous threat from the Romulans. (TNG: "All Good Things...")

In 2369, Amanda Rogers was selected for an internship aboard the Enterprise-D. Doctor Beverly Crusher arranged to have her work in all the major departments aboard the ship while she was there in order to help her get an idea of what field she was most interested in. (TNG: "True Q")

After Captain Edward Jellico took command of the Enterprise-D the same year, he made major changes in every department on the ship. (TNG: "Chain of Command, Part I")

Later that year, Nella Daren became the new department head of stellar sciences aboard the Enterprise-D. In her new role, she was very forthright in her requests for ship resources, such as access to the main sensor array, even as other departments were waiting to use the array. (TNG: "Lessons")

In 2370, after Riker was informed of a fluctuation in the ship's power systems, he authorized a diagnostic while taking the task of informing other departments about the issue upon himself. (TNG: "Genesis")

On USS Voyager, at least thirteen departments were reported to Chief of security Tuvok every day. (VOY: "Scientific Method")

While Voyager was dealing with a singularity in 2371, Captain Janeway ordered that all departments run complete system diagnostics. (VOY: "Parallax")

In 2375, while crossing a desolate region named The Void, Voyager first officer Chakotay ordered all departments to create an energy reserve. (VOY: "Night")

Also in 2375, on the eve of commencing Operation Fort Knox, Janeway noted in her captain's log that all departments were standing by. (VOY: "Dark Frontier")

Specific departmentsEdit

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